Introduction:
The trans-activation response DNA-binding protein (TDP-43) is a DNA/RNA-binding protein located in the nucleus. It is involved in regulating alternative splicing processes and transcriptional activities of RNA.1 A common key pathological hallmark in amyotrophic lateral sclerosis (ALS) and frontotemporal dementia (FTD) is the abnormal aggregation and mislocalization of TDP-43 from the nucleus to the cytoplasm.2In vivo studies have shown that the selective reduction of mislocalized TDP-43 from the cytoplasm is linked to reduced toxicity.3 Since there are currently no treatments which reverse ALS and FTD, TDP-43 represents a promising target to develop novel therapeutic options.
Aim:
To identify novel compounds that can aid in the clearance of cytoplasmic TDP-43 and/or translocate TDP-43 from the cytoplasm back into the nucleus.
Methods:
Human neuroblastoma SH-SY5Y cells stably transfected with disease-relevant TDP-43 M337V mutants were treated with 12 compounds (300 nM) for 4 hours. Cells were fractionated into their cytoplasmic and nuclear fractions. Western blots were performed to determine the effect of compounds on TDP-43 translocation and cytoplasmic clearance.
Results and Discussion:
Compounds JD2-180, TC_ELN197 and TC_ELN215 significantly decreased TDP-43 in the cytoplasm as compared to control (p<0.05). TC_ELN215 was the most effective at decreasing cytoplasmic M337V TDP-43 (no treatment vs TC_ELN215= 0.569 ± 0.008 vs 0.393 ± 0.080, n=3, mean ± SD). These three compounds had no effect on nuclear TDP-43 levels. This suggests that these compounds are able to aid in the clearance of toxic cytoplasmic TDP-43, but are unable to translocate cytoplasmic TDP-43 back into the nucleus.
Conclusion:
We have successfully identified three novel compounds that can selectively decrease mislocated cytoplasmic TDP-43 with no effect on TDP-43 in the nucleus. This provides a scaffold for rational design of more efficacious compounds. Further research will investigate other functional activities such as effects on TDP-43 phosphorylation and other disease-related post-translational modifications.