Zinc (Zn) is the second most abundant transition metal ion in humans and is also a potent antimicrobial component of the innate immune response at the host pathogen interface. Bacteria subvert or resist host Zn insults by metal efflux pathways that include cation diffusion facilitator (CDF) proteins. The structural and functional examination of this protein family has been limited, with only the structures of the YiiP proteins from Escherichia coli (PDB 3H90) [1] and Shewanella oneidensis (PDB 5VRF) [2] described to date.
In this work, we have targeted the C-terminal domains of the CzcD proteins from Cupriavidus metallidurans, Pseudomonas aeruginosa and Thermotoga maritima for structural and functional analysis. We have determined the metal binding properties, solution quaternary structures and three-dimensional architectures of these proteins and revealed remarkable similarity in protein structures but significant diversity in the metal-binding properties. Further we have discovered a potential novel mechanism for metal-promoted dimerization for the Cupriavidus metallidurans and Pseudomonas aeruginosa proteins. This poster will present our findings on structural characterisation and metal-binding properties of these CzcD proteins for better understanding of CDF family of proteins.